Additional Complications to Optimizing MAS

• Multiple QTL with different gene actions and epistatic effects

• Multiple traits - Pleiotropic effects

• Epistatic effects with background genome

• Gametic imprinting

• Inbreeding

Overall

EBV = b1QTL1 + b2QTL2 + b3QTL3 . . .+ EBVPolygenic

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